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Archived from the original on February 5, 2009. They reasoned that if a primer pheromones were on the bedding then the sub-dominant's reproductive function should continue to be suppressed. "Observations on the Winter Aggregates of Two Polistine Paper Wasps (Hymenoptera Vespidae Polistinae. 4142 a b c d Howe 2008,. Isaacson, Walter (October 19, 2014).

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